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th Five Year'''' Period[2011BAC02B01]; Chinese Academy of Sciences[KZCX2-YW-223, KZCX2-YW-225]; Technology Development Program Projects of Shandong Province[2008GG20005006, 2008GG3NS0700]; SDSFC[ZR2009CZ008]; CAS/SAFEA; Shandong Provincial Key Laboratory of Ecology and Environment in the Yellow River Delta[2009KFJJ03]">National Science & T 1 Veterinary Sciences 1